EDTA was used as photophosphorylation uncoupler/inhibitor of ATP synthesis. What is EDTA and what is its mode of uncoupling? What is its mode of uncoupling in cholorplast?  I know EDTA is ethylenediaminetetraacetic acid and is a chelating agent. I do not understand how to answer the question on its mode of uncoupling.

Biochemistry
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EDTA was used as photophosphorylation uncoupler/inhibitor of ATP synthesis. What is EDTA and what is its mode of uncoupling? What is its mode of uncoupling in cholorplast? 

I know EDTA is ethylenediaminetetraacetic acid and is a chelating agent. I do not understand how to answer the question on its mode of uncoupling. 

Jagendorf, A.T, and E. Uribe, “ATP formation caused by acid-base transition of spinach chloroplasts, “ Proceedings of the National Academy of Sciences (USA) 55 (1965): 170-177

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EDTA means ethylene diamine tetraacetic acid. EDTA is a divalent ion chelator. EDTA is also used for the inhibition of many proteases that requires divalent metal ions as cofactors. Chloroplast is a double membrane cell organelle in which the process of photosynthesis occurs. 

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