When researchers first discovered that airflow through a bird’s paleopulmonal parabronchi is unidirectional, the question arose as to whether gas exchange is countercurrent, cocurrent, or cross- current. Some ingenious investigators carried out experiments in which they measured the efficiency of gas exchange between air and blood in duck lungs when parabronchial airflow was in its normal direction and when the direction of parabronchial airflow was artificially reversed. The efficiency did not change. How is this evidence against countercurrent and cocurrent gas exchange? How is this evidence for cross-current exchange?

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